Minor Adventure With A Deer

When animals seem "friendly" or "curious" - it's hard for us not to wonder if we're having meaningful relationship. 

I had been scything "thugs" - aggressive plants - that can derail the recovery of biodiversity. It's complicated; let's forget that part for now.

To put it differently, I was working quietly in the ecosystem, chopping down some plants while protecting other dense, beautiful vegetation. The work is compelling and meditative in a fun, relaxing kind of way. Looking up to check a sound, I'm face to face with a young deer.

She was about 10 feet away. Why so close? Whatever, many deer treat us in this comfortable way, when we're focused on something else. 

I sometimes talk with deer as they approach. Not that I think they understand my words, but it seems neighborly to speak in a calm and lighthearted way, and I suppose my words become meditative thinking on another level. 

In this case I said, "Well, thanks for joining me. I suppose you focus so much on these plants that you can't help wonder what I'm doing with them." She kept moving, but not getting closer or farther away. "Are you going to walk a circle around me?"

Indeed, she did walk a circle around me. I went back to work. She ate as she walked. 

I meditated out loud that we were in this together: "You'd be happy if you understood what I'm doing. Without so-called 'stewardship' - this open oak woods would soon be dense with tall goldenrod and the other thugs, none of which you like to eat." It's true. She can't eat most of that we cull. Thuggish plants tend to be toxic to her. So we're on the same side, in that sense.

She doesn't respond verbally, but she speaks with her presence. Indeed, she completes that circle around me. Staying close. Eating and joining in the refreshing shade on this warm day. 

She's in the Shooting Star zone of Somme Woods. This area was dense buckthorn seven years ago. Little sunlight penetrated. There was little food here for deer or anything else. But rather rare and conservative shooting stars survived here in large numbers. So we prioritized this area to cut the brush and restore the sedges, grasses, wildflowers, and shrubs that the buckthorn had shaded out. And of course, we restored fire. 

The plants blooming in the photo above are sweet black-eyed Susan (yellow) and great blue lobelia, but she and I are standing among more than one hundred uncommon species that are here because we cut the buckthorn, controlled the invasive thugs, and planted restorative seeds. She and I both feel good about this place. 

If you know your plants and look hard at these photos you can also identify tall coreopsis, wood reed, Virginia rye grass, and riverbank rye. Because I know this area, I also feel in the presence of Ridell's goldenrod, willow aster, fringed gentian, gayfeather, and the Threatened species Viola labradorica - most of which she'll eat - not too much I hope. 

By this last photo, the doe has traveled a circle and a half around me. It's been good. But I've finished my work in this spot and drift away, finding more thugs to scythe. At some point I notice that she's gone. It's an experience to wonder about. Wild nature is like this. 
 

What Did Trees Look Like?

In natural woodlands of the tallgrass heartland, for thousands of years, trees did not look like what see today.

They were trees of fire – part of a richness that's mostly gone now. But with the ecological restoration of oak woodlands and savannas, they’re coming back. And for them to do so, it may be important for us to understand them – in part so that our sense of “successful management” doesn’t “enhance” or disfigure them in a misguided direction.

 

Only recently have we started burning wooded ecosystems, and their recovery is in an early stage. There were no "nature photographers" to leave us model images. Perhaps artists who saw remnants of nature can help us learn what to look and hope for. The drawing below of a real bur oak is by Adolph Hoeffler in 1852. The tree remembers and reflects aboriginal times. Its character and history are written all over it. The lower branches were burned off long ago.

Notice that, even with a sensitive artist like Hoeffler, the twenty or so other trees in the drawing get increasingly ill-defined as they recede into the distance. They become closer to “standard trees” – like a kid would draw.

 

Hoeffler’s real tree was clearly a savanna tree. Oak woodland trees were probably somewhat different, though also dependent on fire for their nature. It stirs our minds to look at the renderings of ancient real trees. 

 

One early painter of the Native American landscape was George Catlin. He actually lived for years among the tribes and nations to record their likenesses and ways. He did not focus on individual trees, but he documented the landscape. In the 1832 painting below …

… Catlin is shows a Native American with a rifle shooting at some others. The trees do not look all that real. But the natural landscape looks thrillingly real – grassland dominating heights and slopes, scattered savanna trees here and there, little “draws” or “ravines” with varying collections of trees or shrubs, floodplain islands heavily wooded, a prairie edged by a line of trees along the right edge of the river. 

 

In another painting, The subject is “Blackbird’s Grave” – but this time Catlin gets even more gloriously carried away by the landscape.

A detail of this painting …

… includes a rare example of a landscape artist bothering with the flowers. Biodiversity conservationists are thrilled to see them. On this day, it seems rich biodiversity thus revealed itself in the savannas and wood edges. On some other day, the prairie likely would have been bejeweled too. Once again, the trees look wrong, with ridiculously thin trunks, but their placement on the landscape looks like nature.

 

The rest of this post focuses on paintings by Albert Bierstadt, from the mid 1800s: 

 

Here we’re in a woodland. There are few shrubs or small trees. Early travelers described such open woods as the rule in the tallgrass region. The woodland oaks were said to reproduce only sporadically.

 

Prairie and savanna at the edge of a woodland. The outermost tree is dead, likely killed by fire. The next tree has all lower limbs burned off. The next four trees have lower branches re-growing. Below them is an apparent hazel ruff. For more on that, see the next painting. 

 

In the above detail from a larger (mostly open grassland) scene, two trees are contrasted. To the left is the classic savanna tree with just a few lower limbs. To the right is a spreading tree with a hazel ruff beneath. A “ruff” is a thicket of hazelnut shrubs that gets its name from those high, flat-topped Elizabethan collars. Every fire would burn off the hazel (which is very sensitive to fire but a champion re-sprouter). Then following a couple of years without fire, the flat-topped ruff would grow. Further to the right is another grove, without hazel. Perhaps the oak with the ruff grew in a wet depression that was often spared from fires. 

 

The next painting is from the Yosemite valley, farther west. The same fire-pruned trees are present. 

Bierstadt found all his wild nature well west of the Tallgrass Region, because in his time the vast natural landscapes here were already gone. But similar fires maintained similar trees. The fires in the tallgrass region could burn higher and hotter because the grasses grew higher and thicker thanks to more plentiful rain. Carefully controlled fires today mostly burn lower.  

 

Two grand old survivors. On the right is a tree that has recovered, and suffered, and recovered again, for centuries. 

 

The next Bierstadt painting, "The Emigrants," shows settlers heading west:

The artist paid most attention to people, cattle, wagon, cliff, and sunset. But he loved real trees. The sunset sanctifies a nobility he recognized in the bones of their ecology.  

A last painting is one that sold for $486,400 in 2005 at a Christie’s auction. Bierstadt does not compete with Andy Warhol or Jackson Pollock. His work was less about art and more about nature. We share his reverie. Very likely there were scenes much like this in the tallgrass region, even if Albert Bierstadt found his models farther west. Original nature was beautiful everywhere. In spring, much of flat Illinois was ponded,l as below. 

Those big old trees sure look like oaks. I wish we were able to visit such a place. With good restoration and conservation, some day we could.  

Acknowledgements

George Catlin quit his lawyer job and hurriedly learned to paint in order to document Native American life. He began traveling and painting in 1830 in the company of William Clark (of Lewis and Clark). He went out again on his own four times between 1830 and 1836 – staying carefully beyond "the frontier," where warring between Native Americans and Euro-Americans made travel dangerous. Peaceful villages welcomed him. His documentation of aboriginal America is a great contribution and deserves to be better known.

Adolph Hoeffler is described on the Internet as "an itinerant artist" who came in 1848 from Germany and spent time in the American midwest and in Cuba. 

Albert Bierstadt, a formally trained artist, first travelled west in 1859 with a surveyor to paint wild scenes while they still existed. We are indebted to him. 

Thanks to Christos Economou and Eriko Kojima for proofing and edits. 

The Unexpected Discovery of a Surviving High-Quality Oak Woodland

I woke, as if from a dream (a bad dream). When I saw the photos, I doubted by eyes. The images and plant list suggested a far richer woodland than I’d ever seen - but which I had long looked for, far and wide. 

But Yikes! It had recently been bulldozed, in part. Yet – more important – the treasure mostly still existed in this world. Botanist Dan Carter called the site "jaw-droopingly intact." Conservationist Eriko Kojima, on seeing it for the first time, was so blown-away she declared that she "wouldn't sleep for a week." 

 

Oak woods had seemed dull. Compared to prairies, savannas, or fens – these days the woods excite few natural areas conservationists. The word biodiversity is currently hot; it invokes the tropical rain forest, the prairie white-fringed orchid, coral reefs - but not woods.

 

For context, consider the re-discovery of the prairies and savannas. Only in recent decades did prairie impinge on public consciousness, when John Curtis, Aldo Leopold, May Watts, Robert Betz and a few others opened our eyes. (See Endnote 1: The Re-discovery of Nature.) Now we know that original prairie remnants are extremely rare and rich. If you’ve never had the experience, it’s worth learning to identify the plants and to perceive the structure so you can truly see one. An original prairie is strikingly superior in plant and animal biodiversity to degraded remnants or restorations. We’re learning to care for and expand remnants, but a reconstruction is nothing like an original. 

The discovery of the savanna was unexpected and yet more recent. Even in the 1970s, the Illinois Natural Areas Inventory essentially didn't find them, in part because they're rarer than the prairies and in part because no one had yet learned what to look for. (A good start at belatedly recognizing the savanna emerged from the 1994 North American Conference on Savannas and Barrens.)

But now for true remnant woodlands. Did woods of high quality still exist - especially bur oak woods, the tree that requires the most sun, according to Curtis? Can we study them only through restoration? But now the "discovery" and recognition of the quality of the little bur and white oak Army Lake site goes down into history? How big a discovery is it? 

Individual endangered species have the advantage of being somewhat protected by laws, regulations, and standard practices. But – rarer and more important – a very high quality woodland ecosystem is not so protected. (See Endnote 2. Learning from Army Lake.)

 

One of the photos that seemed marvelously rich for a woodland. Largely high-quality and rare plants make up this turf, including yellow star-grass, robin’s-plantain, forked aster, wood pea, veiny pea, wood betony, low false bindweed, sky-blue aster, wood rush, Penn sedge, running savanna sedge, and blue-eyed grass.
(2016 photo by Dan Carter)

Most woodlands today, if they have any quality at all, impress people who care by their dense displays of common spring ephemeral flora. In contrast, Army Lake Woodland has a dense, diverse, rare flora throughout spring, summer, and fall. That it's principally not ephemeral. Because of grazing or excessive shade at some point, most woodlands have lost most high-quality plants - especially those of summer and fall - as well as the other biota (animals, fungi, bacteria, etc.) associated with them.  (See Endnote 3.)

Few heard of Dan's 2016 discovery the time. Further documentation and analysis was needed. People are busy. The site was supposedly secure in the ownership of the Wisconsin Department of Natural Resources (WDNR). But then came the bulldozers.

“A gut punch ... a failure ... a hill worth dying on.”

I personally heard about the site in 2022 when Christos Economou sent me a link to a remarkable 2020 blog post by the perceptive Dan Carter. I didn't know the man. Later, in response to inquiries, Dan expressed two regrets: 1) as a scientist, he regretted that he wrote that post when he was emotional with anger at the destruction; 2) as a person, he wished he'd been more outspoken in defending the site from development, though it would likely have cost him his job. Many eco-experts work for public agencies, or otherwise get funded by them, or just need to cooperate with them. They speak calmly. When the damage came, it was worse than Dan or his colleagues expected. Dan called it "a gut punch." In retrospect a battle to save this woods would have been "a hill worth dying on." (See Endnote 4: Politics.)

Central to what makes a Grade A prairie remnant striking is the density and diversity of rare plants. When Dan Carter was asked to evaluate Army Lake Woodland as a potential site for a boat launch, he found a woodland as rich as a remnant prairie. (See Endnote 5. Plant Species List.) The plants are important in themselves, but they're also the best indicators of full biodiversity that includes the invertebrates, fungi, bacteria and other biota that are principally associated with true, rich remnants because those organisms rely on the plants. . 

 

Army Lake Woodland had a substantially more diverse and high-quality woodland community than is known anywhere else in southeastern Wisconsin (perhaps anywhere in Wisconsin or northern Illinois?). So, how could it get bulldozed by a conservation agency, when that boat launch could have been put on many other spots on the shore of Army Lake? Recommendations were not followed. Insufficient advocacy and public concern emerged. About a third of this best woodland remnant was destroyed. That's the bad part. But, again, two-thirds of this gem and quite a bit more restorable remnant still exist. 

Next Steps 

In the conservation of small remnants, there are typically three values and associated goals:

1. Save, restore, and study what's there.

2. Expand the size of the remnant, if possible, as small plant and animal populations are vulnerable.

3. Use seed and knowledge to restore larger areas. 

1. Save, restore, and study

An anticipated program of restoration management is crucial, as increasing shade and invasives threaten more and more quality. The site has to date received no invasives control and no prescribed burns. WDNR staff resources are stretched thin and deserve higher appropriations. But volunteer stewards could carry a lot of the load on this small site, if properly facilitated and empowered. Needed stewardship would likely lead to additional species emerging, knowledge, and insights. 

2. Expand

The high-quality remnant biota could be restored into poorer-quality adjacent areas. A first step that could start any time would be invasives control. Crown vetch has recently been discovered along the boat launch access road. Many dangerous invasives are there in small numbers. They should go soon. Some native species now cast damaging amounts of shade, in the absence of fire. Regular controlled burns are a must. Many managers would recommend annual burns of half or more of such a site, at least while it's in the "intensive care ward" stage.  

3. Use seed and knowledge to restore larger areas. 

Can-do management and restoration of Army Lake Woodland would benefit other nearby remnants (and needy oak woodlands throughout the midwest), in part by what could be learned here in the next few years. Also, many nearby areas would benefit from seed from Army Lake, especially once it has recovered overall quality.    

Most of the diversity on the island survives where over-growth by brush has been limited, perhaps due to the intense competition of diverse grasses and wildflowers.

 

If you’re tempted to take a look at this site, I don’t blame you. But please don’t trample such a tiny, vulnerable, precious remnant. You can pretty well observe what’s important through the fence (that was erected to protect it from boat launch and beach traffic) and on the unfenced half of the island. If you have a serious reason to study the flora and can’t restrain yourself from slipping through the fence, a little fishermen’s trail runs from the fence to the east side of the island. Perhaps some people could walk carefully on that? 
 
Additional views of the rich flora of Army Lake Woodland are below: 
 

Blooming in the center of this photo is the formerly common pale vetchling with its cream-colored blossoms. Also in bloom here are wood betony (yellow), shooting star (white), and wild geranium (pink). 

In bloom, Carolina vetch, blue-eyed grass, and wood rush. In bud: false dandelion. Prominent foliage: shooting star, bastard toadflax, and wood betony.

Blooming: yellow star-grass and robin’s-plantain. In bud upper right: Maryland sanicle. Identifiable by their foliage: bastard toadflax, woodland milkweed, NewJersey tea, northern hawkweed, showy goldenrod, white lettuce, false Solomon’s-seal, sedges, a white oak seedling, and bracken fern. 

Note on bracken and soils: Many ecologists these days expect bracken on sandy soils, but Army Lake Woodland soil, where exposed by bulldozing, is not sandy. Sand savannas and woodland remnants are much more common than on rich soils, perhaps because invasives take over more slowly there. Soils exposed by the bulldozing at Army Lake are not sandy. The biodiversity of rich soils survived here for other reasons.

 

Viewed through the fence that now somewhat protects the woods: alumroot, shooting star, blue-eyed grass, wild geranium, and pale vetchling. In seed on right is common dandelion; it's here because the site gets random trampling stress, which should be minimized. 

 

Endnotes

Endnote 1. What is original nature?

For most of human history, the concept of "nature" was a foggy one, at best. More often nature was something poets said "Oh my!" about in flowery language. More recently, as real nature was slipping away, it needed to be understood and defined.

The seminal Illinois Natural Area Inventory (INAI) of the 1970s was considered by many the most authoritative theoretical and practical approach to defining and conserving true nature in its various forms. Soon, as a result, Illinois protected most of its surviving high quality prairie remnants. They were priceless ecological heritage that had gone unrecognized in part because it had long vanished from productive lands over most of the U.S. and, arguably, the human-occupied world. But the Inventory did much less well on the fire-dependent oak communities. In the 1980s, conservationists figured out how to recognize savanna remnants and begin restoring them to high quality. Only in 2011 did oak woodlands get formally recognized as a category separate from maple forest. Conservation and restoration of oak woodlands is much less advanced than for prairies and savannas. 

Like Illinois, Wisconsin has been in the forefront of natural areas conservation, thanks to John Curtis, Aldo Leopold, The Prairie Enthusiasts, and strong communities of professional and volunteer conservationists. As with the savannas in Illinois, the long overdue discovery of Army Lake Woodland provides insights that suggest how to similarly upgrade biodiversity conservation for oak woods - quality recognition, prioritization, and restoration. 

Endnote 2. Learning from Army Lake

This site is now being used by the Wisconsin Department of Natural Resources and others to develop new oak woodland assessment tools. This blog hopes to report on that effort and other Army lake learning, when there's something to report. 

Braving a light rain in June 2022, participants in improving tools for woodland evaluation included, clockwise from the foreground (back view, in red) Ryan O’Conner (Inventory Coordinator, WDNR), Dan Carter (The Prairie Enthusiasts), Brian Miner (Southeast Wisconsin Land Steward, The Nature Conservancy), Matt Zine (Habitat and Management Specialist, WDNR), and Pete Duerkop (District Ecologist, WDNR). 

Endnote 3. Why does this tiny gem still exist?

The site is only about 1.6 acres and was formerly a semi-island. If this find is as important as it seems, the likely reason is that a tamarack swamp separated it from adjacent uplands, and for that reason it escaped the heavy grazing that degraded the biodiversity of most woodlands. The intervening marsh is narrow enough and flammable enough that prehistoric fires could well have spread across or jumped it. But the site has not been known to burn in recent years. The dense diversity may have helped slow the invasion by brush. Higher-quality natural area are somewhat more resistant to invasion by species that would degrade them. 

Endnote 4: Politics

Both professionals and volunteers struggle with the balance between saying too much (thus making enemies) and saying too little (thus being ignored). When Dan Carter assessed Army Lake Woods for the boat launch, his job was with the Southeastern Wisconsin Regional Planning Commission. In his later post on what happened, he wrote:

"I contacted some local stakeholders and advised them of a public meeting (listening session or some such thing) about the launch, warning about potential impacts to something precious and irreplaceable, and I promptly got an upset phone call from the property manager (though I appreciate him not going straight over my head like the Milwaukee County Parks Natural Areas Coordinator did a few years later). In hindsight, I kind of wish he had called my boss. It might have freed me up to more strongly oppose what was about to happen."

"What happened at Army Lake wasn’t just wrong, it was insane. Really, the forked aster was the only thing there with protected status that might have helped it, but that’s also insane, because communities support rare species; it’s not the other way around."

As Dan points out, the kind of public support that might have made the stakes clearer to decision makers – perhaps to a point that Dan would not have felt his job in jeopardy – was lacking. A strong constituency for Army Lake Woods’ biodiversity might, in fact, have allowed conservationists to feel empowered to look for creative potential solutions. A key question is, how do we caring conservationists build such constituency? 

 

Public agencies are necessarily political. That's not a bad thing. "Politics" is another way of saying "democracy." Compromises and trade-offs have to be made. In challenging cases, the responsibility lies not just on the agency employees but on the whole conservation community. At one point decades ago, the entire Illinois Nature Preserves Commission staff got fired (its budget zeroed out) in part because of failures to compromise. A wise and valiant participant in that struggle, Jerry Paulson, the day-to-day director at that time, wrote a comment on this post (see below) that ended: "Public employees who speak-up to protect nature are true heroes! But, they can't be expected to do it all alone. Every state needs a Friends of Illinois Nature Preserves or The Prairie Enthusiasts to advocate for nature." 

Endnote 5: Army Lake Woodland Plant List

Assembled by Dan Carter as part of a "timed meander" assessment. According to the rules of this exercise, the list includes all the plants that can be identified in a brief survey; the investigator stops when they fail to find a new species in five minutes. There are likely many other species present. (A few additional common names were added for this post.) 

Acer saccharum	hard maple, sugar maple (an invader?)
Agrimonia gryposepala	common agrimony, tall hairy agrimony
Amelanchier laevis	Allegheny serviceberry, Allegheny shadblow
Amphicarpaea bracteata	American hog-peanut, hog-peanut
Anemone americana	round-lobed hepatica
Anemone quinquefolia	nightcaps, wood anemone
Antennaria parlinii	Parlin's pussy-toes, plantain pussy-toes
Apocynum cannabinum	hemp-dogbane, Indian hemp
Aquilegia canadensis	Canadian columbine, red columbine, wild columbine
Aralia nudicaulis	wild sarsaparilla
Asclepias exaltata	poke milkweed, tall milkweed
Aureolaria grandiflora	large-flowered yellow false foxglove
Berberis thunbergii	Japanese barberry  (an invader)
Calystegia spithamaea	low bindweed, low false bindweed
Carduus nutans	musk thistle, nodding thistle  (an invader)
Carex cephalophora	oval-headed sedge, short-headed bracted sedge
Carex pensylvanica	common oak sedge, Pennsylvania sedge
Carex siccata	dry-spiked sedge, hay sedge, hillside sedge, running savanna sedge
Carex stricta	tussock sedge
Carya ovata	shagbark hickory, shellbark hickory
Ceanothus americanus	New Jersey tea, red-root
Celastrus scandens	American bittersweet, climbing bittersweet
Cerastium fontanum	common mouse-ear chickweed  (an invader)
Comandra umbellata	bastard-toadflax, false toadflax
Cornus foemina	gray dogwood, northern swamp dogwood, panicled dogwood
Cornus rugosa	round-leaved dogwood
Cornus sericea	red osier dogwood
Dactylis glomerata	orchard grass  (an invader)
Danthonia spicata	poverty danthonia, poverty grass, poverty oat grass
Diervilla lonicera	northern bush-honeysuckle
Elaeagnus umbellata	autumn olive  (an invader)
Equisetum arvense	common horsetail, field horsetail
Erigeron pulchellus	Robin's-plantain
Euphorbia corollata	flowering spurge
Eurybia furcata	forked aster
Festuca saximontana	Rocky Mountain fescue
Fragaria virginiana	thick-leaved wild strawberry, Virginia strawberry,
Frangula alnus	European alder buckthorn, glossy buckthorn
Fraxinus americana	white ash  (an invader?)
Galium boreale	northern bedstraw
Galium concinnum	pretty bedstraw, shining bedstraw
Gaylussacia baccata	Huckleberry
Gentianella quinquefolia	ague-weed, stiff gentian
Geranium maculatum	Crane's-bill, spotted geranium, wild geranium
Helianthus hirsutus	hairy sunflower, oblong sunflower, rough sunflower

Heuchera richardsonii	prairie alumroot, Richardson's alumroot
Hieracium caespitosum	field hawkweed, yellow king-devil  (an invader)
Hieracium umbellatum	narrow-leaved hawkweed, northern hawkweed
Ilex verticillata	common winterberry
Juniperus virginiana	eastern red-cedar  (an invader?)
Krigia biflora	false-dandelion, orange dwarf-dandelion, two-flowered Cynthia
Lactuca canadensis	Canada lettuce, tall lettuce, tall wild lettuce, w
Larix laricina	larch, tamarack
Lathyrus ochroleucus	cream pea-vine, pale vetchling, white pea
Lathyrus venosus	veiny pea
Lonicera dioica	limber honeysuckle, mountain honeysuckle, red hone
Lonicera X bella	Bell's honeysuckle, showy bush honeysuckle  (an invader)
Luzula multiflora	common wood rush
Maianthemum racemosum	false Solomon's-seal, false spikenard, large false Solomon's-seal
Medicago lupulina	black medick  (an invader)
Melilotus albus	white sweet-clover  (an invader)
Melilotus officinalis	yellow sweet-clover  (an invader)
Ostrya virginiana	eastern hop-hornbeam, ironwood  (an invader?)
Parthenocissus inserta	grape woodbine
Pedicularis canadensis	Canadian lousewort, forest lousewort, wood-betony
Phryma leptostachya	American lop-seed
Picea glauca	white spruce
Plantago major	broad-leaved plantain, common plantain  (an invader)
Poa compressa	Canada bluegrass, wiregrass  (an invader)
Podophyllum peltatum	May-apple, wild mandrake
Polemonium reptans	Greek-valerian, spreading Jacob's-ladder
Populus grandidentata	big-tooth aspen, large-toothed aspen
Potentilla simplex	common cinquefoil, old-field five-fingers
Prenanthes alba	lion's-foot, rattlesnake-root, white-lettuce
Primula meadia	eastern shooting-star, pride-of-Ohio
Prunus serotina	wild black cherry
Prunus virginiana	chokecherry
Pteridium aquilinum	bracken, bracken fern
Quercus alba	white oak
Quercus rubra	northern red oak
Ranunculus abortivus	little-leaf buttercup, small-flowered buttercup
Ranunculus fascicularis	early buttercup, thick-root buttercup
Rhamnus cathartica	common buckthorn, European buckthorn  (an invader)
Rosa carolina	Carolina rose, pasture rose
Rosa multiflora	multiflora rose  (an invader)
Sanicula marilandica	black snakeroot, Maryland sanicle
Sisyrinchium albidum	common blue-eyed grass
Solidago speciosa	showy goldenrod
Solidago ulmifolia	elm-leaved goldenrod
Symphyotrichum laeve	smooth blue aster
Symphyotrichum oolentangiense	sky-blue aster
Symphyotrichum urophyllum	arrow-leaved aster, white arrowleaf aster

Taraxacum officinale	common dandelion  (an invader)
Toxicodendron rydbergii	Rydberg's poison-ivy, western poison-ivy  (an invader?)
Trifolium pratense	red clover  (an invader)
Vaccinium myrtilloides	velvet-leaved blueberry
Viburnum opulus	European cranberry-bush  (an invader)
Viburnum rafinesquianum	arrow-wood, downy arrow-wood
Vicia caroliniana	Carolina vetch, pale vetch, wood vetch
Viola sororia	door-yard violet, common blue violet, hairy wood violet
Vitis riparia	frost grape, river bank grape

 

More references, if you're interested

A fine presentation on savannas and woodlands by Dan Carter 

In that presentation, Dan offers the summary below from Veldman et al. It summarizes characteristics of ancient grassland ecosystems from a paper that focuses principally on the tropics. But it offers principles that may apply to our less-well-studied remnants as well. 

Three of the many posts on this blog that explore similar questions in Illinois remnants and restoration:

https://woodsandprairie.blogspot.com/2022/06/hypoxis-experiment-succeeds-in-ecosystem.html

https://woodsandprairie.blogspot.com/2020/10/a-celebration-of-vestal-grove-study.html

https://woodsandprairie.blogspot.com/2018/02/the-somme-prairie-grove-experiment.html

A similar but more objective account of Army Lake Woods - with some additional information - can be found here.

Acknowledgements

This post was written by Stephen Packard. Credit for the original discovery of the significance of Army Lake Woodland goes to Dan Carter. Dan is now with The Prairie Enthusiasts, a highly-respected organization that focuses volunteer and professional resources on stewardship and advocacy for biodiversity. 

Thanks for proofing and edits: to an early version to Dan Carter, and for this version to Matt Evans and Christos Economou.